A model

I've been moving closer to being able to model the converse pleiotopy idea using an individual based simulation.  Since the question is interwoven with the evolution of parental care, I'll probably model that first, and then impose a phenotype of the offspring that "uses up" the investment by the parent.  I should be able to use a two gene model.  One gene will control parental care.  The other gene will control the offspring phenotype, which will be a twofold, pleiotropic phenotype, one will be adult fitness and the other will be the juvenile fitness cost of the parent to rear it.